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Subclass Copepoda
Crustaceans of the subclass Copepoda are found in both marine and freshwater habitats. Globally, there are c. 13,000 species in 10 orders. Almost 350 species (87 genera, 17 families, 6 orders) are known from North American fresh waters (Boxshall & Defaye, 2008).
Copepods are minute crustaceans in the μm-mm size range. The three tagmata of the typical crustacean body plan are recognizable, although the "head" is actually a cephalothorax (cephalosome) because 1-2 thoracic segments are fused to it. The thorax bears biramous appendages, and the abdomen has only a pair of caudal rami. Although trunk segmentation is generally visible, the cylindrical body has a main point of articulation dividing the anterior metasome from the posterior urosome. The shape of the body tagmata in combination with the length of the antennules can be used to distinguish the free-living copepod orders.
Copepods in general are sexually dimorphic, with modifications of the appendages in males for grasping the females and female appendages modified to carry eggs. Development is indirect through a nauplius larval phase.
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Bibliography
- Smith, I.M., D.R. Cook & B.P. Smith. 2010. Water mites (Hydrachnidiae) and other arachnids. [in] J.H. Thorp & A.P. Covich (eds.). Ecology and Classification of North American Freshwater Invertebrates, 3rd edition: 485-586.
- Reid, J.W. & C.E. Williamson. 2010. Copepoda. [in] J.H. Thorp & A.P. Covich (eds.). Ecology and Classification of North American Freshwater Invertebrates, 3rd edition: 829-899.
- Covich, A.P., J.H. Thorp & D.C. Rogers. 2010. Introduction to the Subphylum Crustacea. [in] J.H. Thorp & A.P. Covich (eds.). Ecology and Classification of North American Freshwater Invertebrates, 3rd edition: 695-723.
- Forró, L. N.M. Korovchinsky, A.A. Kotov & A. Petrusek. 2008. Global diversity of cladocerans (Cladocera; Crustacea) in freshwater. Hydrobiologia 595: 177-184. [Available online]
- Boxshall, G.A. & D. Defaye. 2008. Global diversity of copepods (Crustacea: Copepoda) in freshwater. Hydrobiologia 595: 195-207. [Available online]
- Smith, D.G. 2001. Copepoda. Pennak's Freshwater Invertebrates of the United States, 4th edition: 489-517.
- Underhill, J.C., W.D. Schmid & D.E. Gilbertson. 1982. Taxonomic Keys to the Common Animals of the North Central States, exclusive of the parasitic worms, terrestrial insects, and birds, 4th edition. 205 pp.
- Wilson, M.S. & H.C. Yeatman. 1959. Free-Living Copepoda. [in] W.T. Edmondson (ed.). Ward & Whipple Fresh-Water Biology, 2nd edition: 735-861.
- Pennak, R.W. 1940. Ecology of the microscopic Metazoa inhabiting the sandy beaches of some Wisconsin lakes. Ecological Monographs 10(4): 537-615. [Available online]
Web Pages
Classification of Freshwater TaxaOnly taxa found in North America are listed. Formatting indicates if they are known (to us) from North America, the midwestern states or Wisconsin. |
Kingdom Animalia [details] Subkingdom Eumetazoa [details] clade Bilateria [details] clade Protostomia [details] Superphylum Ecdysozoa [details] Phylum Arthropoda [details] Subphylum Crustacea [details] Class Maxillopoda [details]
|
. . . . |
| Family Centropagidae [details] |
. . . . . |
| Genus Limnocalanus [details] |
. . . . . . |
| Species L. macrurus (+ L. grimaldii), L. johanseni |
. . . . . |
| Genus Osphranticum [details] |
. . . . . . |
| Species O. labronectum [detail] |
. . . . . |
| Genus Sinocalanus [details] |
. . . . . . |
| Species S. doerrii [detail] |
. . . . |
| Family Diaptomidae [details] |
. . . . . |
| Genus Aglaodiaptomus [details] |
. . . . . . |
| Species A. clavipes, A. leptopus, A. stagnalis, A. atomicus, A. clavipoides, ..., etc. |
. . . . . |
| Genus Leptodiaptomus (+ Eutrichodiaptomus) [details] |
. . . . . . |
| Species L. pribilofensis, L. ashlandi, L. minutus, L. sicilis, L. siciloides, ..., etc. |
. . . . . |
| Genus Onychodiaptomus [details] |
. . . . . . |
| Species O. birgei, O. sanguineus, O. hesperus, O. louisianensis, O. virginiensis |
. . . . . |
| Genus Skistodiaptomus [details] |
. . . . . . |
| Species S. oregonensis, S. pallidus, S. pygmaeus, S. reighardi, S. bogalusensis, ..., etc. |
. . . . . |
| Genus Arctodiaptomus (+ Pararctodiaptomus) [details] |
. . . . . . |
| Species A. dorsalis (+ A. dampfi), A. arapahoensis, A. bacillifer, A. floridanus, A. kurilensis, A. novosibiricus, A. saltillinus |
. . . . . |
| Genus Diaptomus (+ Pelorodiaptomus) [details] |
. . . . . . |
| Species D. castor, D. glacialis |
. . . . . |
| Genus Hesperodiaptomus [details] |
. . . . . . |
| Species H. eiseni, H. arcticus, H. augustaensis, H. breweri, H. caducus, ..., etc. |
. . . . . |
| Genus Acanthodiaptomus [details] |
. . . . . . |
| Species A. denticornis [detail] |
. . . . . |
| Genus Eudiaptomus [details] |
. . . . . . |
| Species E. gracilis, E. yukonensis |
. . . . . |
| Genus Mastigodiaptomus [details] |
. . . . . . |
| Species M. albuquerquensis, M. nesus, M. texensis |
. . . . . |
| Genus Mixodiaptomus [details] |
. . . . . . |
| Species M. theeli [detail] |
. . . . . |
| Genus Nordodiaptomus [details] |
. . . . . . |
| Species N. alaskaensis [detail] |
. . . . . |
| Genus Sinodiaptomus [details] |
. . . . . . |
| Species S. sarsi [detail] |
. . . . . |
| Genus Epischura [details] |
. . . . . . |
| Species E. lacustris, E. fluviatilis, E. massachusettsensis, E. nevadensis, E. nordenskioeldi |
. . . . . |
| Genus Eurytemora [details] |
. . . . . . |
| Species E. affinis, E. americana, E. arctica, E. bilobata, E. canadensis, ..., etc. |
. . . . . |
| Genus Heterocope [details] |
. . . . . . |
| Species H. appendiculata, H. septentrionalis |
. . . . |
| Family Aetideidae (+ Senecellidae) [details] |
. . . . . |
| Genus Senecella [details] |
. . . . . . |
| Species S. calanoides [detail] |
. . . . |
| Family Acartiidae [details] |
. . . . . |
| Genus Acartiella [details] |
. . . . . . |
| Species A. sinensis [detail] |
. . . . . |
| Genus Acartia [detail] |
. . . . |
| Family Clausocalanidae (+ Pseudocalanidae) [details] |
. . . . . |
| Genus Drepanopus [details] |
. . . . . . |
| Species D. bungei [detail] |
. . . . |
| Family Pseudodiaptomidae [details] |
. . . . . |
| Genus Pseudodiaptomus [details] |
. . . . . . |
| Species P. euryhalinus, P. forbesi, P. inopinus, P. marinus, P. pelagicus |
. . . . |
| Family Tortanidae [details] |
. . . . . . |
| Species T. dextrilobatus [detail] |
. . . . |
| Family Cyclopidae [details] |
. . . . . |
| Subfamily Cyclopinae [details] |
. . . . . . |
| Genus Acanthocyclops [details] |
. . . . . . . |
| Species A. vernalis, A. capillatus, A. einslei, A. venustoides, A. brevispinosus, ..., etc. |
. . . . . . |
| Genus Diacyclops [details] |
. . . . . . . |
| Species D. bicuspidatus, D. thomasi, D. albus, D. bisetosus, D. languidoides, ..., etc. |
. . . . . . |
| Genus Mesocyclops [details] |
. . . . . . . |
| Species M. edax, M. americanus, M. leuckarti, M. longisetus, M. ogunnus, ..., etc. |
. . . . . . |
| Genus Microcyclops (+ Cryptocyclops) [details] |
. . . . . . . |
| Species M. bicolor, M. dubitabilis, M. pumilis, M. rubellus, M. varicans |
. . . . . . |
| Genus Cyclops [details] |
. . . . . . . |
| Species C. strenuus, C. canadensis, C. columbianus, C. kolensis (+ C. alaskaensis), C. scutifer, C. vicinus |
. . . . . . |
| Genus Megacyclops [details] |
. . . . . . . |
| Species M. latipes (+ M. ingens), M. viridis, M. donnaldsoni, M. gigas, M. magnus |
. . . . . . |
| Genus Orthocyclops [details] |
. . . . . . . |
| Species O. modestus [detail] |
. . . . . . |
| Genus Thermocyclops [details] |
. . . . . . . |
| Species T. dybowskii, T. oithonoides, T. crassus, T. inversus, T. parvus, T. tenuis |
. . . . . . |
| Genus Apocyclops [details] |
. . . . . . . |
| Species A. barri, A. dengizicus, A. dimorphus, A. panamensis, A. spartinus |
. . . . . . |
| Genus Bryocyclops [details] |
. . . . . . . |
| Species B. muscicola [detail] |
. . . . . . |
| Genus Itocyclops [details] |
. . . . . . . |
| Species I. ishidai, I. pulaskii, I. yezoensis |
. . . . . . |
| Genus Metacyclops [details] |
. . . . . . . |
| Species M. cushae, M. gracilis |
. . . . . . |
| Genus Rheocyclops [details] |
. . . . . . . |
| Species R. carolinianus, R. hatchiensis, R. indiana, R. talladega, R. virginianus |
. . . . . . |
| Genus Stolonicyclops [details] |
. . . . . . . |
| Species S. heggiensis [detail] |
. . . . . |
| Subfamily Eucyclopinae [details] |
. . . . . . |
| Genus Macrocyclops [details] |
. . . . . . . |
| Species M. ater, M. albidus (+ M. bistriatus), M. fuscus |
. . . . . . |
| Genus Paracyclops [details] |
. . . . . . . |
| Species P. yeatmani, P. fimbriatus, P. affinis, P. bromeliacola, P. canadensis, ..., etc. |
. . . . . . |
| Genus Ectocyclops [details] |
. . . . . . . |
| Species E. phaleratus, E. polyspinosus |
. . . . . . |
| Genus Eucyclops [details] |
. . . . . . . |
| Species E. agilis (+ E. montanus), E. bondi, E. borealis, E. conrowae, E. elegans, ..., etc. |
. . . . . . |
| Genus Tropocyclops [details] |
. . . . . . . |
| Species T. prasinus, T. extensus |
. . . . . . |
| Genus Homocyclops [details] |
. . . . . . . |
| Species H. ater [detail] |
. . . . . |
| Subfamily Halicyclopinae [details] |
. . . . . . |
| Genus Halicyclops [details] |
. . . . . . . |
| Species H. bowmani, H. clarkei, H. coulli, H. fosteri, H. hurlberti, ..., etc. |
. . . . |
| Family Lernaeidae [details] |
. . . . . |
| Genus Lernaea [detail] |
. . . . |
| Family Oithonidae [details] |
. . . . . |
| Genus Limnoithona [details] |
. . . . . . |
| Species L. sinensis, L. tetraspina |
. . . |
| Order Harpacticoida [details] |
. . . . |
| Family Parastenocarididae [details] |
. . . . . |
| Genus Parastenocaris [details] |
. . . . . . |
| Species P. brevipes (+ P. wilsoni, + P. starretti), P. lacustris, P. palmerae, P. trichelata |
. . . . . |
| Genus Horstkurtcaris [details] |
. . . . . . |
| Species H. delamarei [detail] |
. . . . |
| Family Phyllognathopodidae [details] |
. . . . . |
| Genus Phyllognathopus [details] |
. . . . . . |
| Species P. viguieri [detail] |
. . . . . |
| Genus Nitokra (+ Nitocra) [details] |
. . . . . . |
| Species N. hibernica, N. bdellurae, N. bisetosa, N. evergladensis, N. lacustris, N. spinipes, N. typica |
. . . . . |
| Genus Nitocrella [details] |
. . . . . . |
| Species N. incerta, N. subterranea |
. . . . . |
| Genus Novanitocrella [details] |
. . . . . . |
| Species N. aestuarina [detail] |
. . . . . |
| Genus Psammonitocrella [details] |
. . . . . . |
| Species P. boultoni [detail] |
. . . . . |
| Genus Stygonitocrella [details] |
. . . . . . |
| Species S. sequoyahi [detail] |
. . . . . |
| Genus Nitocrellopsis [detail] |
. . . . |
| Family Canthocamptidae [details] |
. . . . . |
| Genus Attheyella (+ Neomrazekiella, + Mrazekiella) [details] |
. . . . . . |
| Species A. americana, A. pilosa, A. alaskaensis, A. carolinensis, A. dentata, ..., etc. |
. . . . . |
| Genus Bryocamptus (+ Arcticocamptus, + Limocamptus) [details] |
. . . . . . |
| Species B. hiemalis, B. hutchinsoni, B. minutus, B. nivalis, B. pygmaeus, ..., etc. |
. . . . . |
| Genus Canthocamptus [details] |
. . . . . . |
| Species C. assimilis, C. robertcokeri, C. staphylinoides, C. staphylinus, C. vagus, C. oregonensis, C. sinuus |
. . . . . |
| Genus Cletocamptus (+ Marshia) [details] |
. . . . . . |
| Species C. albuquerquensis, C. brevicaudatus, C. deborahdexterae, C. fourchensis, C. stimpsoni |
. . . . . |
| Genus Elaphoidella [details] |
. . . . . . |
| Species E. bidens, E. subgracilis, E. amabilis, E. californica, E. fluviusherbae, ..., etc. |
. . . . . |
| Genus Gulcamptus [details] |
. . . . . . |
| Species G. huronensis, G. laurentiacus |
. . . . . |
| Genus Heteropsyllus [details] |
. . . . . . |
| Species H. nunni, H. pseudonunni |
. . . . . |
| Genus Maraenobiotus [details] |
. . . . . . |
| Species M. insignipes, M. brucei, M. canadensis, M. husmanni, M. subterraneus |
. . . . . . |
| Species M. alaskana, M. lilljeborgii, M. mexicana, M. pygmaea, M. rapiens, M. wolskii |
. . . . . . |
| Species M. affinis, M. cristata, M. duthiei, M. hudsoni, M. mrazeki, M. laurentica (+ M. americana), M. virginiana |
. . . . . |
| Genus Epactophanes [details] |
. . . . . . |
| Species E. richardi [detail] |
. . . . . |
| Genus Pesceus (+ Paracamptus) [details] |
. . . . . . |
| Species P. reductus, P. reggiae |
. . . . . |
| Genus Stenocaris [details] |
. . . . . . |
| Species S. minor [detail] |
. . . . |
| Family Harpacticidae [details] |
. . . . . |
| Genus Harpacticella [details] |
. . . . . . |
| Species H. paradoxa [detail] |
. . . . . |
| Genus Harpacticus [details] |
. . . . . . |
| Species H. chelifer, H. gracilis |
. . . . . |
| Genus Tigriopus [details] |
. . . . . . |
| Species T. californicus (+ T. triangulus) [detail] |
. . . . |
| Family Laophontidae [details] |
. . . . . |
| Genus Onychocamptus [details] |
. . . . . . |
| Species O. mohammed, O. talipes |
. . . . . |
| Genus Heterolaophonte [details] |
. . . . . . |
| Species H. stroemii [detail] |
. . . . . |
| Genus Pseudonychocamptus [details] |
. . . . . . |
| Species P. proximus [detail] |
. . . . . |
| Genus Microlaophonte [detail] |
. . . . |
| Family Miraciidae (+ Diosaccidae) [details] |
. . . . . |
| Genus Schizopera [details] |
. . . . . . |
| Species S. borutzkyi, S. knabeni, S. tobae |
. . . . |
| Family Tachidiidae [details] |
. . . . . |
| Genus Microarthridion [details] |
. . . . . . |
| Species M. littorale [detail] |
. . . . . . |
| Species G. incisipes [detail] |
. . . . . |
| Genus Tachidius [details] |
. . . . . . |
| Species T. discipes, T. spitzbergensis |
. . . . |
| Family Canuellidae [details] |
. . . . . . |
| Species C. canadensis [detail] |
. . . . |
| Family Chappuisiidae (+ Uchidastygacaridae) [details] |
. . . . . |
| Genus Chappuisius, Uchidastygacarus |
. . . . |
| Family Cletodidae [details] |
. . . . . . |
| Species K. breviarticulatum [detail] |
. . . . . |
| Genus Limnocletodes [details] |
. . . . . . |
| Species L. behningi [detail] |
. . . . |
| Family Dactylopusiidae [details] |
. . . . . |
| Genus Paradactylopodia [details] |
. . . . . . |
| Species P. brevicornis [detail] |
. . . . |
| Family Darcythompsoniidae [details] |
. . . . . |
| Genus Leptocaris [details] |
. . . . . . |
| Species L. brevicornis, L. kunzi, L. mangalis |
. . . . |
| Family Ectinosomatidae [details] |
. . . . . |
| Genus Pseudobradya [detail] |
. . . . . . |
| Species M. jousseaumei [detail] |
. . . . |
| Family Nannopodidae (+ Huntemanniidae) [details] |
. . . . . |
| Genus Huntemannia [details] |
. . . . . . |
| Species H. jadensis, H. lacustris |
. . . . . . |
| Species N. palustris [detail] |
. . . |
| Order Siphonostomatoida [details] |
. . . . |
| Family Lernaeopodidae [details] |
. . . . . |
| Genus Salmincola, Achtheres, Cauloxenus |
. . . . . |
| Genus Lepeophtheirus [detail] |
. . . . |
| Family Dichelesthiidae [detail] |
. . . |
| Order Poecilostomatoida [details] |
. . . . |
| Family Ergasilidae [details] |
. . . . . |
| Genus Neoergasilus [details] |
. . . . . . |
| Species N. japonicus [detail] |
. . . . . |
| Genus Ergasilus [details] |
. . . . . . |
| Species E. chautauquaensis [detail] |
. . . . . |
| Genus Thersitina [detail] |
. . . . |
| Family Scaeogelyellidae [details] |
. . . . . |
| Genus Scaeogelyella [details] |
. . . . . . |
| Species S. caroliniana [detail] |
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